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Family BREGMACERIDAE GILL 1872�@ƒTƒCƒEƒI‰È
Bregmacerids (ƒTƒCƒEƒI‰È‹›Ží)
Bregmacerotids (ƒTƒCƒEƒI‰È‹›Ží)
Codlets (ƒTƒCƒEƒI—Þ)
Unicorn-Cods (ƒTƒCƒEƒI—Þ)


Genus Bregmacerina^♀ DANIL'CHENKO 1957�@ƒuƒŒƒOƒ}ƒPƒŠƒi‘® (�â–Å)

�ŸƒuƒŒƒOƒ}ƒPƒŠƒi�EƒAƒ“ƒeƒBƒNƒA (�â–Å)
�@Bregmacerina antiqua (Smirnov 1936)
�@�@�@syn.Motella tricirrata antiqua Smirnov 1936
�@^*1936�w„Q„�„q„� „ƒ„u„r„u„‚„€-„{„p„r„{„p„x„ƒ„{„€„s„€ „€„|„y„s„€„ˆ„u„~„p, „X„u„‚„~„€„‚„u„‰„u„~„ƒ„{„y„z „‚„p„z„€„~�F„B„|„p„t„y„}„y„‚ „P„u„„„‚„€„r„y„‰ „R„}„y„‚„~„€„r�x�B
�@^*1957�w„N„€„r„�„z „‚„€„t „ƒ„u„}„u„z„ƒ„„„r„p Bregmacerotidae in „M„p„„„u„‚„y„p„|„� „{ „€„ƒ„~„€„r„p„} „�„p„|„u„€„~„„„€„|„€„s„y„y 1�F„P„p„r„u„| „C„u„€„‚„s„y„u„r„y„‰ „D„p„~„y„|„Ž„‰„u„~„{„€�x�B
�@^*1960�w„K„€„ƒ„„„y„ƒ„„„�„u „‚„�„q„� „}„p„z„{„€„�„ƒ„{„y„‡ „€„„„|„€„w„u„~„y„z „K„p„r„{„p„x„p�F„P„p„r„u„| „C„u„€„‚„s„y„u„r„y„‰ „D„p„~„y„|„Ž„‰„u„~„{„€�x�B

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Genus Bregmaceros^♂ THOMPSON 1840�@ƒTƒCƒEƒI‘®
�@�@�@�¦�{Genus Asthenurus^♂ TICKELL 1865 (Type species : Asthenurus atripinnis Tickell 1865)�B
�@�@�@�¦�{Genus Calloptilum^⚲ RICHARDSON 1845 (Type species : Calloptilum mirum Richardson 1845)�B
�@�@�@�¦�‚Genus Calloptilum^⚲ HUTTON 1873 [Moridae]�B
�@�@�@�¦�{Genus Podopteryx^♂ SAUVAGE 1880 (Type species : Podopteryx albyi Sauvage 1880)�B
�@�@�@�¦�‚Genus Podopteryx^♀ SELYS 1871 [Megapodagrionidae, Zygoptera, Insecta]�B
�@�@�@�¦�‚Genus Podopteryx^♀ SHAROV 1971 [Sharovipterygidae, Protorosauria, Reptilia]�B
�@�@�@�¦�{Genus Pseudeleginus^♂ SAUVAGE 1873 (Type species : Pseudeleginus albyi Sauvage 1873)�B
�@�@�@�¦�{Genus Problennius^♂ PRIEM 1908 (Type species : Problennius filamentosus Priem 1908)�B


�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒAƒ‹ƒrƒC (�â–Å)
�@Bregmaceros albyi (Sauvage 1881)
�@�@�@syn.Podopteryx albyi Sauvage 1881 non Sauvage 1873
�@^*1881�wNouvelles Recherches sur les Poissons Fossiles Decouvertes par M.Alby a Licata, en Sicile in Annales des Sciences Géologiques 11�FHenri-Émile Sauvage�x�B
�@^*2004�wAdditions à l'Ichthyofaune Tortonienne du Bassin de Ierapetra, Crète Orientale, Grèce�FJean Gaudant�x�B
�@^*2015�wFish-Otoliths from the Marine-Brackish Water Transition from the Middle Miocene of the Belgrade Area, Serbia�FWerner W.Schwarzhans, Katarina Z.Bradić-Milinović & Ljupko Rundić�x�B

�Ÿ
�@Bregmaceros anchovius Ho, Endo, Lee & Chu 2020
�@^*2020�wBregmaceros anchovia sp.nov., a New Codlet Species from the Western Pacific Ocean [Gadiformes, Bregmacerotidae]�FHsuan-Ching Ho, Hiromitsu Endo, Chia-Lien Lee & Tah-Wei Chu�x�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒAƒ“ƒeƒBƒNƒEƒX (�â–Å)
�@Bregmaceros antiquus Schwarzhans 1980
�@^*1980�wDie Tertiäre Teleosteer-Fauna Neuseelands, Rekonstruiert Anhand von Otolithen�FWerner W.Schwarzhans�x�B
�@^*1984�wNew Zealand Cenozoic Teleost Fishes, Represented by Fossil Otoliths in Phylum Chordata, Lancelets, Fishes, Amphibians, Reptiles, Birds, Mammals�FCarolyn M.King, Clive D.Roberts, Ben D.Bell, R.Ewan Fordyce, Robert S.Nicoll, Trevor H.Worthy, Chris D.Paulin, Rod A.Hitchmough, Ian W.Keyes, Alan N.Baker, Andrew L.Stewart, Norton Hiller, Robert M.McDowall, Richard N.Holdaway, Robin P. McPhee, Werner W.Schwarzhans, Alan James Drummond Tennyson, Seabourne Rust & Ian Macadie�x�B
�@‚P‚X‚W‚O”N”Å‚ª“ƈíŒê�E‚P‚X‚W‚S”N”Å‚ª‰pŒê�B

�Ÿ
�@Bregmaceros arabicus


�ŸƒZƒCƒˆƒEƒTƒCƒEƒI
�@Bregmaceros atlanticus Goode & Bean 1886
�@^*1896�wA Check-List of the Fishes and Fish-Like Vertebrates of North and Middle America 1895�FDavid Starr Jordan & Barton Warren Evermann�x�B
�@^*1896�wOceanic Ichthyology, Based upon a Study of the Deep-Sea Fishes of the Atlantic Basin�FGeorge Brown Goode & Tarleton Hoffman Bean�x�B
�@^*1898�wThe Fishes of North and Middle America 3, a Descriptive Catalogue of the Species of Fish-Like Vertebrates Found in the Waters of North America, North of the Isthmus of Panama�FDavid Starr Jordan & Barton Warren Evermann�x�B

�Ÿ
�@Bregmaceros bathymaster Jordan & Bollman 1890
�@�@�@syn.Bregmaceros longipes Garman 1899
�@^*1890�wScientific Results of Explorations by the US Fish Commission Steamer Albatross 4, Descriptions of New Species of Fishes Collected at the Galapagos Islands and Along the Coast of the United States of Colombia, 1887-88 1889�FDavid Starr Jordan & Charles Harvey Bollman�x�B
�@^*1899�wReports on an Exploration off the West Coasts of Mexico, Central and South America and off the Galapagos Islands in Charge of Alexander Agassiz by the U.S.Fish Commission Steamer Albatross, During 1891, Lieut.Commander Z.L.Tanner, U.S.N., Commanding 16, the Fishes�FSamuel Walton Garman�x�B
�@^*1899�wReports on an Exploration off the West Coasts of Mexico, Central and South America and off the Galapagos Islands in Charge of Alexander Agassiz by the U.S.Fish Commission Steamer Albatross, During 1891, Lieut.Commander Z.L.Tanner, U.S.N., Commanding 16, the Fishes Plates�FSamuel Walton Garman�x�B
�@"ƒRƒbƒhƒŒƒbƒg"�BIUCN Least Concern ver.3.1�B


�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�Eƒ{ƒXƒjƒAƒXƒL (�â–Å)
�@Bregmaceros bosniaski (Sauvage 1881)
�@�@�@syn.Podopteryx bosniaski Sauvage 1881 non Sauvage 1873
�@^*1881�wNouvelles Recherches sur les Poissons Fossiles Decouvertes par M.Alby a Licata, en Sicile in Annales des Sciences Géologiques 11�FHenri-Émile Sauvage�x�B

�Ÿ
�@Bregmaceros cantori
�@"ƒXƒgƒ‰ƒCƒvƒRƒbƒh"�B

�Ÿ
�@Bregmaceros cayorus

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�Eƒ_ƒjƒNƒX (�â–Å)
�@Bregmaceros danicus Schwarzhans & Nielsen 2023
�@^*2023�wFish Otoliths from the Bathyal Eocene Lillebalt Clay Formation of Denmark�FWerner W.Schwarzhans & Kent Albin Nielsen�x�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒfƒNƒ‰ƒXƒW (�â–Å)
�@Bregmaceros deklaszi Schwarzhans 2010
�@^*2010�wOtoliths from the Miocene of West Africa, Primarily from the Mandorové Formation of Gabon�FWerner W.Schwarzhans�x�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒtƒFƒ‹ƒPƒŠ (�â–Å)
�@Bregmaceros felkeri Schwarzhans 2007
�@^*2007�wThe Otoliths from the Middle Eocene of Osteroden Near Bramsche, North-Western Germany�FWerner W.Schwarzhans�x�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒtƒBƒ‰ƒ�ƒ“ƒgƒXƒX (�â–Å)
�@Bregmaceros filamentosus (Priem 1908)
�@�@�@syn.Problennius filamentosus Priem 1908
�@^*1908�wPoissons Fossiles de Perse in Revue Critique de Paleozoologie 3�FFernand Priem�x�B
�@^*1908�wPoissons Fossiles de Perse, Mission de Morgan�FFernand Priem�x�B

�Ÿ
�@Bregmaceros houdei
�@"ƒnƒEƒTƒCƒEƒI"�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒqƒuƒŠƒhƒDƒX (�â–Å)
�@Bregmaceros hybridus Schwarzhans 2010
�@^*2010�wOtoliths from the Miocene of West Africa, Primarily from the Mandorové Formation of Gabon�FWerner W.Schwarzhans�x�B
�@^*2014�wOtoliths from the Middle Miocene, Serravallian of the Karaman Basin, Turkey�FWerner W.Schwarzhans�x�B


�ŸƒTƒCƒEƒI
�@Bregmaceros japonicus (Tanaka 1911) Jordan & Hubbs 1925
�@�@�@syn.Bregmaceros atlanticus japonicus Tanaka 1911
�@^*1908-1911�wDescription of Eight New Species of Fishes from Japan 1908�FShigeho Tanaka�x�B
�@^*1913�wA Catalogue of the Fishes of Japan�FDavid Starr Jordan, Shigeho Tanaka & John Otterbein Snyder�x�B
�@^*1913-1914�wRecord of the Fishes Obtained in Japan in 1911�FDavid Starr Jordan & William Francis Thompson�x�B
�@^*1922-1925�wRecord of Fishes Obtained by David Starr Jordan in Japan 1922�FDavid Starr Jordan & Carl Leavitt Hubbs�x�B
�@^*2023�wEarly Pliocene Otolith Assemblages from the Outer-Shelf Environment Reveal the Establishment of Mesopelagic Fish Fauna Over 3 Million Years Ago in South-Western Taiwan�FChien-Hsiang Lin, Siao-Man Wu, Chia-Yen Lin & Chi-Wei Chien�x�B

�Ÿ
�@Bregmaceros lanceolatus
�@‹D�…ˆæ�i“ü�«�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�Eƒ‹ƒGƒ‹ƒŠƒ“ƒQƒlƒ“ƒVƒX (�â–Å)
�@Bregmaceros luellingenensis Schwarzhans & Wienrich ex Wienrich 2009
�@^*2009�wDie Fauna des Marinen Miozäns von Kevelaer, Niederrhein 5, Vertebraten�FGünther Wienrich�x�B


�Ÿƒ~ƒiƒ~ƒTƒCƒEƒI
�@Bregmaceros mcclellandi Thompson 1840
�@�@�@syn.Asthenurus atripinnis Tickell 1865
�@�@�@�@�@Bregmaceros albyi (Sauvage 1873)
�@�@�@�@�@Bregmaceros atripinnis (Tickell 1865) Day 1878
�@�@�@�@�@Bregmaceros catulus Schubert 1908
�@�@�@�@�@Bregmaceros macclellandi Jordan 1897�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Bregmaceros macclellandii Alcock 1893�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Bregmaceros maclellandi Hardenberg 1931�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Calloptilum mirum Richardson 1845
�@�@�@�@�@Otolithus (Genus Xenodermichthys) catulus Schubert 1908
�@�@�@�@�@Pseudeleginus albyi Sauvage 1873 non Sauvage 1881
�@�@�@�@�@Xenodermichthys catulus (Schubert 1908)
�@^*1843-1845�wIchthyology 1+2+3 1844-1845 in the Zoology of the Voyage of the H.M.S. Sulphur, under the Command of Captain Sir Edward Belcher R.N., C.B., F.R.G.S., etc, During the Years 1836-42�FJohn Richardson�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma, and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1893�wNatural History Notes from H.M. Indian Marine Survey Steamer, Investigator, Commander C.F.Oldham, R.N., Commanding Series 2 Number 9, an Account of the Deep Sea Collection Made During the Season of 1892-93�FAlfred William Alcock�x�B
�@^*1896�wA Check-List of the Fishes and Fish-Like Vertebrates of North and Middle America 1895�FDavid Starr Jordan & Barton Warren Evermann�x�B
�@^*1897�wNotes on Fishes, Little Known or New to Science 1896�FDavid Starr Jordan�x�B
�@^*1898�wThe Fishes of North and Middle America 3, a Descriptive Catalogue of the Species of Fish-Like Vertebrates Found in the Waters of North America, North of the Isthmus of Panama�FDavid Starr Jordan & Barton Warren Evermann�x�B
�@^*1899�wA Descriptive Catalogue of the Indian Deep-Sea Fishes in the Indian Museum, Being a Revised Account of the Deep-Sea Fishes Collected by the Royal Indian Marine Survey Ship Investigator�FAlfred William Alcock�x�B
�@^*1904�wRésultats du Voyage du S.Y. Belgica en 1897-1898-1899, Sous le Commandement de A. de Gerlache de Gomery, Rapports Scientifiques Publiés aux Frais du Gouvernement Belge, Sous la Direction de la Commission de la Belgica, Zoologie, Poissons�FLouis Dollo�x�B
�@^*1904�wThe Fishes of Panama Bay�FDavid Starr Jordan & Edwin Chapin Starks�x�B
�@^*1905�wOn a Collection of Fishes from the Inland Sea of Japan, Made by Mr. R.Gordon Smith�FCharles Tate Regan�x�B
�@^*1908�wDie Fischotolithen des Pausramer Mergels�FRichard Johann Schubert�x�B
�@^*1912�wDie Fischfauna der Schliermergel von Bingia Fargeri, bei Fangario in Sardinien�FRichard Johann Schubert�x�B
�@^*1931�wThe Fish-Fauna of the Rokan Mouth�FJohann Dietrich Franz Hardenberg�x�B
�@^*2019�wPaleogene Fish Otoliths [Teleostei] from the Subsilesian and Ždánice units in Moravia�FRostislav Brzobohatý & Miroslav Bubík�x�B
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@‹D�…ˆæ�i“ü�«�B

Bregmaceros atripinnis=Bregmaceros mcclellandi

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�Eƒ~ƒkƒgƒDƒX (�â–Å)
�@Bregmaceros minutus Stinton 1958
�@^*1958�wFish Otoliths from the Tertiary Strata of Victoria 1957�FFrederick Charles Stinton�x�B

�Ÿ
�@Bregmaceros moseri Harold & Baltzegar 2023
�@^*2023�wA New Species of Bregmaceros [Gadiformes, Bregmacerotidae] from the Eastern Pacific Ocean, with Comments on B.atlanticus and B.japonicus�FAntony S.Harold & D.Andrew Baltzegar�x�B

�Ÿƒgƒ„ƒ}ƒTƒCƒEƒI
�@Bregmaceros nectabanus Whitley 1941
�@�@�@syn.Bregmaceros nactabanus Koeda, Hata, Yamada & Motomura 2018�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@^*2018�w�•’ª‚ ‚½‚鎭Ž™“‡‚ÌŠC�A“à”V‰Y‹™�`‚É�…—g‚°‚³‚ê‚é‹›‚½‚¿�F�¬Ž}Œ\‘¾�^”¨�°—Ë�^ŽR“cŽç•F�^–{‘º�_”V�x�B
�@^*2021�w한국 제주도에서 날개멸과, 대구목 어류, Bregmaceros nectabanus 의첫 출현 및 북서태평양 날개멸과 8종의 종검색�F장서하�^김진구�x�B
�@^*2023�wEarly Pliocene Otolith Assemblages from the Outer-Shelf Environment Reveal the Establishment of Mesopelagic Fish Fauna Over 3 Million Years Ago in South-Western Taiwan�FChien-Hsiang Lin, Siao-Man Wu, Chia-Yen Lin & Chi-Wei Chien�x�B

�ŸƒNƒ�ƒnƒ‰ƒTƒCƒEƒI
�@Bregmaceros neonectabanus

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�EƒIƒuƒ�ƒ“ƒOƒX (�â–Å)
�@Bregmaceros oblongus Schwarzhans 1977
�@^*1977�wOtolithen aus dem Unteroligozän, Tertiär von Häckelhoven, Kreis Heinsberg, Nordrhein-Westfalen�FWerner W.Schwarzhans�x�B

�Ÿ
�@Bregmaceros pescadorus
�@"‘ä˜pƒTƒCƒEƒI"�B

�Ÿ
�@Bregmaceros pseudolanceolatus Torii, Javonillo & Ozawa, 2004

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�Eƒvƒ�ƒ\ƒ|ƒmƒX (�â–Å)
�@Bregmaceros prosoponos Grenfell 1984
�@^*1984�wEarly Miocene Teleost Otoliths from Parengarenga Harbour, New Zealand�FHugh R.Grenfell�x�B
�@^*2021�wFish Otoliths from the Early Miocene of Chile, a Window into the Evolution of Marine Bony Fishes in the South-East Pacific�FWerner W.Schwarzhans & Sven Nikolaus Nielsen�x�B

�Ÿ
�@Bregmaceros pseudolanceolatus
�@"ƒtƒH�[ƒ‹ƒXƒ‰ƒ“ƒXƒRƒbƒh"�B

�Ÿ
�@Bregmaceros rarisquamosus
�@"ƒIƒIƒ�ƒTƒCƒEƒI"�B

�Ÿ
�@Bregmaceros retrodorsalis Ho, Endo & Chu 2020
�@^*2020�wA New Species of the Codlet Genus Bregmaceros from the Western Pacific Ocean [Gadiformes, Bregmacerotidae]�FHsuan-Ching Ho, Hiromitsu Endo & Tah-Wei Chu�x�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒX�Eƒgƒ�ƒGƒ‹ƒŠ (�â–Å)
�@Bregmaceros troelli Frizzell & Dante 1965
�@^*1965�wOtoliths of Some Early Cenozoic Fishes of the Gulf Coast�FDonald Leslie Frizzell & John H.Dante�x�B
�@^*1999�wIchthyofaunen aus dem Atlantischen Tertiär der USA, Systematik, Paläoäkologie, Biostratigraphie und Paläobiogeographie 1997�FArnold Müller�x�B
�@^*2022�wMiddle and Late Eocene Fish Otoliths from the Eastern and Southern USA�FChien-Hsiang Lin & Dirk Nolf�x�B

�ŸƒCƒ“ƒhƒTƒCƒEƒI
�@Bregmaceros sp.
�@^*2017�wŽ­Ž™“‡‘åŠw‘��‡Œ¤‹†”Ž•¨ŠÙ‚É�Š‘ ‚³‚ê‚Ä‚¢‚éˆÝ“à—e•¨‹›—Þ•W–{�F�¬Ž}Œ\‘¾�^–{‘º�_”V�x�B

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Genus Bregmacerotidarum^⚲�@ƒuƒŒƒOƒ}ƒPƒ�ƒeƒBƒ_ƒ‹ƒ€‘® (�â–Å)�@�¦Otoliths��Ž¨�Î�B

�ŸƒuƒŒƒOƒ}ƒPƒ�ƒeƒBƒ_ƒ‹ƒ€�Eƒ~ƒjƒ€ƒ€ (�â–Å)
�@Bregmacerotidarum minimum (Frost 1934)
�@�@�@syn.Otolithus (Genus Macrurus) minimus Frost 1934
�@^*1934�wOtoliths of Fishes from the Lower Tertiary Formations of Southern England 5�FGeorge Allan Frost�x�B

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Genus Maorigadus^♂ SCHWARZHANS 1980�@ƒ}ƒIƒŠƒKƒhƒDƒX‘® (�â–Å)

�Ÿƒ}ƒIƒŠƒKƒhƒDƒX�EƒAƒiƒ‹ƒLƒNƒX (�â–Å)
�@Maorigadus anarchicus Schwarzhans 1980
�@�@�@syn.Macrouridarum anarchicum (Schwarzhans 1980)
�@^*1980�wDie Tertiäre Teleosteer-Fauna Neuseelands, Rekonstruiert Anhand von Otolithen�FWerner W.Schwarzhans�x�B
�@^*1984�wNew Zealand Cenozoic Teleost Fishes, Represented by Fossil Otoliths in Phylum Chordata, Lancelets, Fishes, Amphibians, Reptiles, Birds, Mammals�FCarolyn M.King, Clive D.Roberts, Ben D.Bell, R.Ewan Fordyce, Robert S.Nicoll, Trevor H.Worthy, Chris D.Paulin, Rod A.Hitchmough, Ian W.Keyes, Alan N.Baker, Andrew L.Stewart, Norton Hiller, Robert M.McDowall, Richard N.Holdaway, Robin P. McPhee, Werner W.Schwarzhans, Alan James Drummond Tennyson, Seabourne Rust & Ian Macadie�x�B
�@‚P‚X‚W‚O”N”Å‚ª“ƈíŒê�E‚P‚X‚W‚S”N”Å‚ª‰pŒê�B

�Ÿƒ}ƒIƒŠƒKƒhƒDƒX�EƒfƒŠƒJƒgƒDƒ‹ƒX (�â–Å)
�@Maorigadus delicatulus Schwarzhans 1980
�@^*1980�wDie Tertiäre Teleosteer-Fauna Neuseelands, Rekonstruiert Anhand von Otolithen�FWerner W.Schwarzhans�x�B

�Ÿƒ}ƒIƒŠƒKƒhƒDƒX�EƒEƒNƒ‰ƒCƒjƒNƒX (�â–Å)
�@Maorigadus ukrainicus Schwarzhans & Bratishko 2011
�@^*2011�wThe Otoliths from the Middle Paleocene of Luzanivka, Cherkasy District, Ukraine�FWerner W.Schwarzhans & Andriy Volodymyrovich Bratishko�x�B

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